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Percichthyidae, the temperate perches, are a family of freshwater found in Australia and southern South America. Formerly placed in the order , they are now placed in the order .

The name Percichthyidae derives from the Latin perca for perch and Ancient Greek ἰχθύς, ichthys for fish.


Evolution
Potential percichthyid fossil remains are known from the of the El Molino Formation of Bolivia; these remains have been described as resembling those of the fossil Percichthys hondoensis from the of . One articulated specimen is known from the Paleocene portion of the formation, but other remains are known from the beds of the formation, indicating that percichthyids could have potential origins. However, the condition of this material is unclear. Another fossil percichthyid, Percichthys lonquimayiensis from , was initially dated to the Paleocene, but further studies have found it to be from the .

Potential percichthyid scales have been recovered from the -aged fossil deposits of in New Zealand. If these are of percichthyids, this suggests they had a wider distribution during the . They also appear to have ranged as far north as the Neotropics during the Oligocene/Miocene, with the fossil genus known from Brazil.

Modern percichthyids have a significant disjunct distribution, being found in Australia and southern South America. All extant members of the group only inhabit freshwater habitats, and it is thus unlikely they ever evolved marine habits. For this reason, it is likely that the modern distribution of percichthyids derives from dispersal over the Antarctic land bridge between South America and Australia, which existed up to the end of the Eocene. This suggests that percichthyids likely inhabited Antarctica as well.


Classification
The temperate perches are closely related to the of North America, with both belonging to the suborder . They were previously also thought to be allied with the temperate basses of the family and the seabasses in the family , but are now thought to not be closely related.

Almost 40 of percichthyids are now recognised, grouped in 11–12 . Most but not all are exclusively freshwater fishes. They are mainly found in , but species are also found in southern South America ( Percichthys).

More recently the Chinese perches have been classified in the separate family while the genus has been found not to be closely related to either that family or the Percichthyidae and has been placed in its own family .

(2025). 9781118342336, Wiley. .
The two species in are found in Australia and were previously placed in Macquaria, but phylogenetic analyses have found them to be the most basal members of the whole order. Abstract

The following 8 genera are classified within the family Percichthyidae:

In addition, two genera are known exclusively from remains:


Species
Australia has the greatest number of percichthyid species, where they are represented by the Australian freshwater cods ( Maccullochella spp.), which are , Mary River cod, eastern freshwater cod, and , by the Australian freshwater blackfishes ( Gadopsis spp.), which are and two-spined blackfish, and by the Australian freshwater perches which are , ( Macquaria spp.), and , and ( Percalates spp.).

Several other Australian freshwater species also sit within the family Percichthyidae, while research using mitochondrial suggests the species of the family Nannopercidae are in reality percichthyids, as well. Australia is unique in having a freshwater fish fauna dominated by percichthyids and allied families/species. This in contrast to Europe and Asia, whose fish faunas are dominated by members of the Cyprinidae carp family. (Australia does not have a single naturally occurring cyprinid species; unfortunately, the illegal introduction of carp has now established the family's presence in Australia.)

A single genus occurs outside Australia, Percichthys in southern South America.

A number of species are or have been important food species; some of these (e.g. the Murray cod, Maccullochella peelii peelii) have become through and river regulation, while others are now to some extent. Some smaller species (e.g. Balston's pygmy perch, Nannatherina balstoni) are popular in .

The extremely rare Bloomfield River cod, Guyu wujalwujalensis, is only found in a short stretch of the Bloomfield River in north .


Timeline
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See also


External links

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